Monday, 6 June 2011

Kildonan, Isle of Arran

Arran has a very varied geology, which has given rise to a rich diversity of shore types. The shores at Kildonan are a mixture of white sandy bays, shingle beaches and basalt dykes (with plenty of rockpools).

Kildonan Port a'Ghille Ghlais raised shore
Raised Beach near Kildonan at Port a'Ghille Ghlais

The luxuriantly vegetated cliffs of the hinterland are a classic example of a post-glacial raised shore, formed when land rose after the melting of glacial ice sheets, whose weight previously kept this land depressed.

Kildonan Tertiary basalt dykes
Tertiary basalt dykes at Kildonan

Basalt dykes and sills form tall, lichen-crusted rock-walls which chop the shore into segments and extend into the sea. These intrusions were formed in the Tertiary when molten magma intruded into pre-existing rock layers, in this case Triassic mudstone (sedimentary rock formed from clay or mud).

When many dykes arise from a single volcanic event they form what is known as a 'dyke swarm', of which Kildonan has one of the best examples in the world.

More in-depth info on Kildonan's geology can be found at these sites:

Coastal Zone Assessment Survey of Port a' Ghille Ghlais to Kildonan (Map 5) produced by SCAPE & Historic Scotland.

Site Report by the Joint Nature Conservation Committee (JNCC)

Pladda Ailsa Craig
Isle of Pladda (left) and the Ailsa Craig (right) from Kildonan
Pladda Ailsa Craig
Isle of Pladda with Ailsa Craig behind

Pladda
Lighthouse on Pladda

The Isle of Pladda is a flat-topped volcanic sill and was formed when magma flowed sideways between pre-existing rock layers. Once cooled, this magma was harder than surrounding rocks and resisted erosion, forming a cliff above softer rocks. It has had a lighthouse since 1790, though the current building dates from around 1830.

Ailsa Craig
Ailsa Craig

The Ailsa Craig (also known as Paddy's Milestone) is a sharply conical volcanic plug composed mostly of microgranite (a very hard, fine-grained granite) which contains the minerals riebeckite and aenigmatite. It has the 3rd largest Gannet colony in the UK and many of these birds can be seen fishing around the coasts of Arran.

Kildonan is a great place to see Common Seals (Phoca vitulina): a large colony is present all year and the seals seem surprisingly tolerant of human presence. As we approached the seals we could hear their snorts, grunts and wails echoing off the green cliffs curving round the bay.

Dog-owners should keep their pets on a leash along this stretch of shore to avoid disturbing the seals, as they haul out very close to the coastal path.

Common Seal Phoca vitulina
Common Seal (Phoca vitulina)
Common Seal Phoca vitulina
Common Seal (Phoca vitulina)

Common Seal Phoca vitulina
Common Seal (Phoca vitulina)

Common Seal Phoca vitulina
Common Seal (Phoca vitulina)
Common Seal Phoca vitulina
Common Seal (Phoca vitulina)

Common Seals (Phoca vitulina) can be distinguished from Grey Seals (Halichoerus grypus) by the following features:

  • V-shaped nostrils (the Grey Seal has almost parallel nostrils).
  • Rounded head with short snub-nosed muzzle. Pronounced dip between forehead and muzzle (the Grey Seal has a long 'Roman nosed' muzzle and a flat forehead). 
  • Eyes positioned on the front of the face, closer to the nose than the back of the head (Grey Seals have their eyes positioned slightly on the sides of the head, halfway between nose and back of head).
  • Fur is very finely and uniformly mottled with pale outlines/rings around darker grey spots (Grey Seals have more strikingly marked, blotchier coats often with sharply contrasting patterning).
Common Seals are the smaller of the two species, weighing 45-105 kg (males are only slightly bigger than females). Unlike Grey Seals, Common Seals are born without a woolly white fetal coat (called lanugo) as it is shed in the womb.

The Common Seal's scientific name means 'calf-like seal' (Phoca = seal: from Latin which is derived from the Greek phoke, vitulina = calf-like: Latin).

Common Seal Phoca vitulina
Common Seal (Phoca vitulina)

Seals have highly sensitive whiskers known as vibrissae (those on the muzzle are mystacial vibrissae) which are used in the tactile detection and identification of prey.

Common Seal Phoca vitulina
Common Seals (Phoca vitulina)

Seals may look sluggish and ungainly on land but in the water they're amazingly graceful and possess an amazing array of adaptations to their marine habitat including the ability to fully collapse their lungs when diving, the ability to drink seawater and the presence of myoglobin in their muscles (which has a 40% higher oxygen-carrying capacity than haemoglobin).

Common Seal Phoca vitulina
Common Seals (Phoca vitulina)

Some of these Common Seals will be heavily pregnant females and will give birth over the next few weeks (Grey Seals pup at a different time of year: from September til early December).

Common Gull Larus canus first-summer
Common Gull (Larus canus) first-summer

We found an Oystercatcher's nest on the upper shore, it contained four cream eggs marked with blackish speckles and scribbles - the parent quickly and quietly returned to incubate once we were a few metres away.

A quick list of some of the birds seen today in Kildonan bay: a Curlew, 3+ pairs of Oystercatchers, a Hooded Crow, 2 Gannets, countless Rock Pipits, numerous Common and Herring Gulls, 5 Shelduck, 3 Merganser and a Buzzard.

Common Stork's-bill Erodium cicutarium
Common Stork's-bill (Erodium cicutarium)


Common Stork's-bill is a delicate pink-petalled flower related to geraniums, so-named because its long seed pods supposedly resemble storks' bills.

The seeds of this plant each possess a long bristle (known as an awn) which gives them the unusual ability to bury themselves. These awns, although composed of dead cells, are hygroscopically-active: they straighten when wet and coil into corkscrew helices when dry.
Coiling of the awn causes the seed to be fired from the seedpod and the motor action which results from the coiling and uncoiling of the awn causes the seed to move across the ground surface - until it falls into a crevice - and then enables it to drill itself into the soil.

A detailed account of this unusual process can be found here.

Dove's-foot Crane's-bill Geranium molle
Dove's-foot Crane's-bill (Geranium molle)
Hemlock Water Dropwort Oenanthe crocata
Hemlock Water Dropwort (Oenanthe crocata)
Hemlock Water Dropwort Oenanthe crocata
Hemlock Water Dropwort (Oenanthe crocata)
Hemlock Water Dropwort Oenanthe crocata
Hemlock Water Dropwort (Oenanthe crocata)

Hemlock Water Dropwort has leaves which smell appetisingly of Celery and bear a strong likeness to Lovage, however the whole plant (particularly the tuberous root) is deadly poisonous. It contains oenanthetoxin which affects the central nervous system, causing severe convulsions and in many cases, death.

How to Identify Hemlock Water Dropwort:

  • The whole plant is entirely hairless.
  • Leaves are 3-4 pinnate, toothed, slightly glossy and parsley-scented.
  • Stout stem is grooved.
  • Flowers are white and possess both upper and lower bracts.
3-4 pinnate = leaves are composed of leaflets arranged along an axis, each of which is further subdivided into leaflets which are themselves subdivided (and again for 4-pinnate leaves).

Northern Marsh Orchid Dactylorhiza purpurella
Western Marsh Orchid (Dactylorhiza purpurella)
Northern Marsh Orchid Dactylorhiza purpurella
Western Marsh Orchid (Dactylorhiza purpurella)
Rock Arch
Sea Milkwort Glaux maritima
Sea Milkwort (Glaux maritima)
Sea Milkwort Glaux maritima
Sea Milkwort (Glaux maritima)
Smooth Hawksbeard Crepis capillaris
Smooth Hawksbeard (Crepis capillaris)
Smooth Hawksbeard Crepis capillaris
Smooth Hawksbeard (Crepis capillaris)
Rock Pipit Anthus petrosus
Rock Pipit (Anthus petrosus)
Wren Troglodytes troglodytes
Wren (Troglodytes troglodytes)
Wren Troglodytes troglodytes
Wren (Troglodytes troglodytes)
Wren Troglodytes troglodytes
Wren (Troglodytes troglodytes)
Wren Troglodytes troglodytes
Wren (Troglodytes troglodytes)
Wren Troglodytes troglodytes
Wren (Troglodytes troglodytes)

I followed this little Wren as it hopped over boulders and deftly snatched insects out of the air.

Male Wrens are polygamous: each male builds a series of nests on his territory and tries to entice females to visit them: female Wrens prefer males who have more nests.
A female who is impressed with his collection of 'cock's nests' will mate with him and line a nest with feathers before laying her eggs. The male will then set about attracting another female to another one of his nests (the females incubate the eggs & do most of the young-rearing).

lichens
Lichens - Lecanora rupicola?
lichens
Lichens - Lecanora rupicola?

As they compete for space, these lichens form a beautiful map-like pattern of crackled continents with boldly demarcated borders.

I found them growing on boulders in the supralittoral or 'splash zone' of the upper shore - I think they *might* be Lecanora rupicola.

Orange-tip female Anthocharis cardamines
Orange-tip (Anthocharis cardamines) female

We walked back along the bay, until we reached Kildonan Hotel: to the side of the carpark there is a huge, rusty propeller leaning against the wall of a white-washed boathouse. House Sparrows nest in gaps under the roof.

House Sparrow Passer domesticus male
House Sparrow (Passer domesticus) male
House Sparrow Passer domesticus male
House Sparrow (Passer domesticus) male
House Sparrow Passer domesticus male
House Sparrow (Passer domesticus) male
House Sparrow Passer domesticus male
House Sparrow (Passer domesticus) male
Brown-lipped Banded Snail Cepaea nemoralis hyalozoate yellow
Brown-lipped Banded Snail (Cepaea nemoralis)

The Brown-lipped Banded Snail (Cepaea nemoralis) is one of two Cepaea species found in Britain (the other being Cepaea hortensis - the White-lipped Banded Snail). Due to the extreme polymorphism of shell patterning and colour found in both species, they can only be identified conclusively by dissection. However, most Brown-lipped Snails have a brown lip to their shells.

Studies on the genetics of shell colour and pattern in Cepaea nemoralis (here and here) have demonstrated that brown shell colour is dominant over both pink and yellow, that pink colouration is dominant over yellow, unbanded is dominant to banded patterns, and that darker banding is dominant over paler banding.

Evolutionmegalab.org has excellent photos of the various colour morphs.

My specimen is a yellow, hyalozonate (transparent banded) individual (very faint double banding is visible in the full-size photo). Some shell morphs provide better camouflage in certain habitats - yellow snails are more common in grassland and darker snails predominate in woodland. Shell colour also affects the thermal properties of a shell: lighter (yellow) shells reduce heat absorption.

I collected lots of holey stones from the shingle shore between Port a'Ghille Ghlais and Bennan Head:

piddock holes Hiatella arctica pebble
Pebbles with bivalve (Hiatella arctica?) bore holes
piddock holes Hiatella arctica pebble
Pebbles with bivalve (Hiatella arctica?) bore holes
piddock holes Hiatella arctica pebble
Pebbles with bivalve (Hiatella arctica?) bore holes
piddock holes Hiatella arctica pebble
Pebbles with bivalve (Hiatella arctica?) bore holes
piddock holes Hiatella arctica pebble
Pebbles with bivalve (Hiatella arctica?) bore holes
piddock holes Hiatella arctica pebble
Pebbles with bivalve (Hiatella arctica?) bore holes
piddock holes Hiatella arctica pebble
Pebbles with bivalve (Hiatella arctica?) bore holes
piddock holes Hiatella arctica pebble
Pebbles with bivalve (Hiatella arctica?) bore holes

These smoothly drilled holes, up to 1.5cm in diameter, are the work of bivalve molluscs: most likely the Wrinkled Rock Borer (Hiatella arctica), given the location and size of burrow.

Juveniles attach themselves to substrates using byssus threads and gradually carve burrows for themselves using the valves of their shells to mechanically abrade solid rock.

Thursday, 26 May 2011

Mugdock Country Park, East Dunbartonshire

I joined West Dunbartonshire's Over 50's Walking Group on a trip to Mugdock Country Park where there are ancient 'bluebell woods' and the opportunity to see a variety of warblers!

We walked northwest from Mugdock Country Park Visitor Centre through mixed woodland near the Craigallian & Carbeth Lochs and followed the path along the western edge of Craigallian Loch to the deciduous woodland to the south. Then we took the track leading eastwards to the grounds of Craigend Castle (and the ruins of Craigend Zoo) and returned to the visitor centre.

In the woods northwest of Mugdock Castle we passed an old, rotten tree in which there was a Great Spotted Woodpecker's nesthole - the young could be heard peeping. Soon after, one of the parents was sighted nearby.

Willow Warblers sang from every scrubby tree as we entered a marshy clearing to the east of the Carbeth & Craigallian Lochs. Here we saw a pair of Roe Deer run down the hillside and a Grey Heron which flew over and made an ungracious and unsuccessful attempt to land on top of a tall pine.

Suddenly, a large, long-winged bird of prey glided past us through the clearing in the direction of Carbeth Loch: an Osprey!

Definitely the highlight of the day for me, as I have never seen one of these birds so close to home.

On the path around Craigallian Loch we heard the singing of Sedge Warblers concealed amongst the vegetation: their song is a fast, frenetic (slightly crazy) warble full of harsh cackles and chattering.

Ancient woodland (south of the visitor centre)

In the woods surrounding Craigend Castle we could hear the songs of both Chiffchaffs and Willow Warblers. We also saw a pair of Treecreepers.

Beech tree (Fagus sylvestris)
Distribution transformer on telegraph poles
Bistort (Persicaria bistorta)

Bistort leaves are the main ingredient of Dock pudding (also known as Easter Ledge Pudding), a fried dish traditionally eaten during Lent in Yorkshire (where the World Dock Pudding Competition is held each April in Calderdale), Cumberland and the Lake District.

Dock Pudding Recipes:

Dock Pudding Recipe from BBC Food

Assorted Dock Pudding Recipes from Historicalfoods.com

Vegetarian Dock Pudding Recipe from Voyageofthebattenberg.blogspot.com

Monday, 2 May 2011

Kilpatrick Hills, West/East Dunbartonshire

Another search for the elusive Adders at Burncrooks Reservoir!

I walked with my parents from Duntocher to the Humphrey Reservoir, past Duncolm and the Lily Loch, to Burncrooks Reservoir and on the way home we crossed the Slacks and Little Round Top instead of following the Humphrey Road.

The Humphrey Road and the Slacks afford some brilliant aerial views of the River Clyde:

The River Clyde from the Slacks
The River Clyde from the Slacks

We cut across the moorland (to the left of the Humphrey Road) where we found a Meadow Pipit's nest in the heather: a neat grass-lined cup holding a set of four tiny, chocolate-mottled eggs.

On the path around Duncolm I saw the first Green Hairstreak butterfly of the day: small, rapidly darting, with wings brown on top and irridescent green underneath.
The drystone wall near the Lily Loch was covered with basking Green Hairstreaks - they fluttered erratically in the sunshine and when they landed they orientated their wings sharply in the direction of the sun.

Green Hairstreak (Callophrys rubi)
Looking towards the Lily Loch from the base of Duncolm - the Green Hairstreak's habitat

Hundreds of huge Horse Leeches (Haemopis sanguisuga) could be seen in the Lily Loch's peaty waters (still bare of vegetation), fattening themselves on a glut of tadpoles.

These large leeches are safe to handle as they're unable to bite through human skin - in Britain, only the rare Medicinal Leech (Hirudo medicinalis) is capable of this (it doesn't stop them from sucking onto you though!).

This plump leech is (rather endearingly) still clutching the reed it grabbed onto as it tried to avoid capture:

Horse Leech (Haemopis sanguisuga)
The Lily Loch with Duncolm behind

The Lily Loch is part of the Dumbarton Muir SSSI (Site of Special Scientific Interest) - it's the only oligotrophic (nutrient poor) loch in Dunbartonshire and is surrounded by an area of raised bog.

Burncrooks Reservoir

As we crossed the grassy hill where the river feeds into Burncrooks, a Sparrowhawk flew up from the ground.

We saw a pair of Greylag Geese (Anser anser) feeding at the edge of Burncrooks Reservoir - these are nervous wild birds and hard to get close to, quickly entering the water the minute they see you. I tried to creep closer through the rushes but the geese began to honk with alarm and were soon swimming out of sight.

Greylag Goose (Anser anser)
Greylag Geese (Anser anser)

We searched amongst the dried golden grasses, dark heather, scree and the russet, wiry tangle of old Bracken fronds...but didn't find any Adders.

On the grassy bank of the smallest Burncrooks Dam, we found a glittering Heath Goldsmith beetle (Carabus nitens).

The river which feeds into the Lily Loch, Duncolm behind
The Slacks
Orange-tip (Anthocharis cardamines) female on foodplant - Cuckoo Flower (Cardamine pratensis)
Wild Pansies (Viola tricolor)
Bracken (Pteridium aquilinum)
Little Round Top Wood

Saturday, 30 April 2011

Kilpatrick Hills, West Dunbartonshire


Little Round Top Wood (OS map grid reference - NS4773) is an excellent 'bluebell wood' rarely visited by walkers - I spent the day listening for the songs of arriving warblers and trying to immortalize with photos the ephemeral newly-minted-lushness of spring foliage.

Bluebell (Hyacinthoides non-scripta)
Bluebell (Hyacinthoides non-scripta)
Crab Apple (Malus sylvestris)
Crab Apple (Malus sylvestris)

Cultivated or Domestic Apples (Malus domestica) are mostly descended from the Central Asian Wild Apple (Malus sieversii) which was brought to Europe and later crossed with closely related 'Malus series' apples, including our native Crab Apple (Malus sylvestris).

Cuckoo Flower (Cardamine pratensis)
Cuckoo Flower (Cardamine pratensis)

Cuckoo Flower (Cardamine pratensis)

Blackthorn or Sloe (Prunus spinosus)
crab spider (Xysticus cristatus) female
Dog's Mercury (Mercurialis perennis) in Little Round Top Wood
Dog's Mercury (Mercurialis perennis) in Little Round Top Wood
Dog's Mercury (Mercurialis perennis) in Little Round Top Wood

At this time of year, Dog's Mercury (Mercurialis perennis) forms a thick carpet of fresh green leaves in Little Round Top Wood. Few creatures seem to nibble at this pristine foliage, possibly due to its poisonous nature.
Culpeper's Complete Herbal (published in 1653) warns 'there is not a more fatal plant, native of our country, than this'. Ingestion causes severe haemorrhagic inflammation of the gastrointestinal tract and kidneys.

Ground Ivy (Glechoma hederacea)
Gorse (Ulex europaeus)

I had a look at the steepest part of Little Round Top Wood (above the marsh) which will be overgrown with nettles & bracken, and inaccessible within a month or two.

Hairy St. John's-wort (Hypericum hirsutum) grows on the scrubby slope and Square-stalked St. John's-wort (Hypericum tetrapterum) grows in the marsh below. St. John's-worts, of which there are many species, are better identified by their leaves and stalks than by their golden-yellow flowers (which appear in late June).

Hairy St. John's-wort (Hypericum hirsutum)
Hairy St. John's-wort (Hypericum hirsutum)


Hairy St. John's-wort has a rounded, hairy stem and the leaves are hairy, elliptical in shape and do not possess the black gland-dots which are found on many other Hypericum species.

Wild Pansies (Viola tricolor)